英语翻译They cannot store water,and their reliance on an entirely unexceptional kidney results in a rate of water loss through renal function significantly higher than that of desert rats.through是谓语吗 that of是指前文的描述的rats，

英语翻译They cannot store water,and their reliance on an entirely unexceptional kidney results in a rate of water loss through renal function significantly higher than that of desert rats.through是谓语吗 that of是指前文的描述的rats，
英语翻译
They cannot store water,and their reliance on an entirely unexceptional kidney results in a rate of water loss through renal function significantly higher than that of desert rats.
through是谓语吗 that of是指前文的描述的rats，那为什么用that of来指代呢？

英语翻译They cannot store water,and their reliance on an entirely unexceptional kidney results in a rate of water loss through renal function significantly higher than that of desert rats.through是谓语吗 that of是指前文的描述的rats，
他们不能储存水,由于依赖于非常普通的肾,导致水流失比跳鼠严重.
through不是谓语,results 才是.因为这里有比较级,所以用that 指代